I downloaded your file and opened it up to take a look. The first I looked at was the sprites collision polygon. The entire polygon has been pushed to the left side of the sprite and a little above the base. In your post you said,
[quote:35ol991x]The sprites seem to be correct in their bounding box in the image editor...
I am not sure what you looked at but, the collision polygon is the problem. Open the image editor and select the bottom item in the left side menu. It looks like a bunch of dots connected by lines and will say "Set collision polygon" when you hover over it.